Professional Documents
Culture Documents
Canada
A catalog of
Nearctic
Chironomidae
A catalog of Catalogue des
Nearctic Chironomidae
Chironomidae delardgion
ndarctique
D.R. Oliver and M.E. Dillon D.R. Oliver et M.E. Dillon
Biosystematics Research Centre Centre de recherches biosyst6matiques
Ottawa,Ontario Ottawa (Ontario)
K1A 0C6 K1A 0C6
and et
P.S. Cranston P.S. Cranston
and
Commonwealth Scientific Organisation de la recherche
Research
Industrial scientifique et industrielle du
Entomology
Organization, Commonwealth, Entomologie
2601
Canberra ACT Canberra ACT 2601
Australia Australie
Price subject to change without notic Prix sujet i changemenl sans pr6avis
Cover Couverture
Adult male of Chirnmw pfumm (Linnaeus) M6le adulte du Chironomw plumm (Linnaeus)
noter que ce ne sont pas tous les groupes qui Chironomus ffiannulatus Macquart,
se sont fait assigner des tribus et des sous- 1826: 202.-Germany.
genres). La premidre fois qu'un nom valide Cricotopus exilis Johannsen, 1905:
apparait, il est imprim6 en caractires gras 255.-N.Y.
prds de la marge de gauche de chaque
colonne. Les taxons valides qui ont 6t Le nom de I'espdce est suivi d'un compte
signalds a tort comme de la rdgion n6arctique rendu synoptique de la rdpartition dans la
sont aussi numdrs le long de la marge de rdgion ndarctique indiqude par le territoire, la
gauche, mais en italique, comme le sont province ou I'Etat, dans le sens nord-ouest a
d'ailleurs toutes les autres listes de noms, nord-est et sud-ouest d sud-est, sauf pour le
valides ou non. Groenland. L'apparition de l'espdce dans
Les citations pour les noms de groupes de d'autres rdgions zoog,ographiques est
genres se prsentent comme suit : 6numr6e ensuite et sdpar6e par un point-
virgule. Quand l'apparition d'une espdce est
Genus NANOCLADIUS Kieffer indiqude dans une r6gion du Mexique situde
Nanocladius Kieffer. 1913d: 31. prds de la division impr6cise entre les rdgions
Type-species, vitellinus Kieffer narctique et notropicale, elle est indiqude
(orig. des.). comme observable tant au Mexique que dans
Microcricotopas Thienemann & la rdgion n6otropicale. La citation binomiale
Harnisch, 7932: 737. originale apparait ensuite, suivie des sous-
Type-species, Cricotopus especes ou varidtds, puis des synonymes (s'il y
albicornis Goetghebuer a lieu), qui sont mentionnds sous leur forme
(Goetghebuer in Goetghebuer & originale accompagns du nom de I'auteur, de
Lenz,1944; 114) = Chironomus la date, de la page et de l'emplacement type.
bicolor Zetterstedt. Les emplacements types pour le Canada et
les Etats-Unis sont indiquds selon les
Subgenus Nanocladius Kieffer frontidres des provinces, des territoires ou
des Etats; tous les autres suivent les
La citation originale se trouve sur la frontidres des pays ou encore des grandes
deuxi0me ligne, en retrait, suivie des 169ions g6ographiques si celui-ci n'est pas
synonymes (s'il y a lieu), eux aussi en retrait. connu.
Le nom du genre est suivi du nom de
I'auteur original, de la date, de la page, du ABREVIATIONS
nom de I'espdce type et de la nature de la Les abrdviations utilises sont les
fixation du type. L'esp0ce type originale est suivantes :
mentionnee, mais le genre est omis s'il est G6n6ralitds
identique d celui dont elle est le type. Si le &- et
nom de l'espOce type originale est par contre E.-
e., est, de l'est, orientale
un synonyme mineur, il apparait i la fin emend.- corrigd
aprds un symbole d'6galit6 (=). Seuls les I., Is.- ile, iles
auteurs de fixation de type ultdrieure sont ICZN.- Commission internationale de
mentionnds. Une grande partie des nomenclature zoologique
informations sur la fixation des types et les misident.- erreur d'identification
synonymes sont tirdes de Ashe (1983). Cette mon.- monotypie
publication contient des renseignements nam.- nom
d6taill6s sur la synonymie g6n6rique, les n. nam.- nouveau nom
espoces types et les probldmes de nom. nud.- nom seul
nomenclature. Les citations des noms des orig. des.- dsignation originale
sous-genres suivent celles des genres, sauf s'il preocc.- d6jd appliqud
s'agit de rdpdtition (dans ce cas, elles sont s., S.- sud, du sud, mdridional
alors omises). se.,Se.- sud-est, du sud-est
Les noms des groupes d'espdces se sub. mon.- monotypie ulterieure
presentent comme suit : subg.- sous-genre
syn.- synonyme, synonymie
triannulatus. Sask., Ont. to N.Y., s. to var.- varidtd
Ky. and Del.; PA. w.- ouest, de I'ouest, occidental
NEARCTIC CHIRONOMIDAE
Subfamily PODONOMINAE
Genus TRICIIOTANYPUS Kieffer
Tribe BOREOCHLINI Trichotanypus Kieffer, 1906a:. 42.
Type-species, Tanypus posticalis
Genus BOREOCHLUS Edwards Lundbeck (mon.).
Boreochlus Edwards in Edwards &
Thienemann, t938: I52. Type-species, alaskensis. Alas.
thienemanni Edwards (orig. des.). Trichotanypus alaskensis Brundin, 1966:
315.-Alas.
gracilistylus. Alas., Alta., Wash., Wyo. foliaceus. Alas.
Boreochlus gracilistylus Brundin, 1966: Trichotanypus foliaceus Wirth & Sublette,
304.-Alta. 1970: 343.-Alas.
persimilis. Alas. to Qu., Calif. to Va. hanseni. Wyo.
Trichotanypus persimilis Johannsen, 1926: Trichotanypus hanseni Wirth & Sublette,
99.-N.Y. 1970: 347.-Wyo.
NEARCTIC CHIRONOMIDAE
scutellata. N.W.T. s. to Fla., Grld.; PA.; NT; festivellus. Man., Ont.; PA.
AU. Chironomus festivellus Kieffer, 1906a: 18
Corynoneura scutellata Winnertz, 1846: (n. nam. for Chironomus festivus
13.-Germany. Meigen, 1830:. 252, Preocc. SaY,
taris. Ohio, N.Y., s. to Miss. & Fla. 1823).-Europe.
Corynoneura /aris Roback, 1957a: flavocinctus. Man.; PA.
61.-N.J. Tricho cl adius fl av o c inc tus Kieffer,
1924b: 92.-Germany.
fugax. Ont., N.Y., Ky., N.C., s. to Miss. &
Genus CRICOTOPUS Wulp Fla.
Cricotopus Wulp 1874: 132. Type-species, Orthocladius fugax Johannsen, 1905:
Chironomus tibialis Meigen (Coquillett 279.-N.Y.
1910b: 528). fuscus. Ont.; PA.
Trichocladius fuscus Kieffer, 1909:
Subgenus CRICOTOPUS Wulp 46.-Germany.
gelidus. N.W.T., Grld.; PA.
abanus. B.C., Man. Trichocladius gelidus Kieffer, 1922b:
Cricotopus abana Curran, 1929: 1.-Man. 18.-Novaya Zemlya.
absurdus. N.Y., Ohio, N. Mex. infuscatus. Ont., S. Dak., Ill., Ohio, Pa., N.
Orthocladius absurdus Johannsen. 190-5: Mex.
277.-N.v. Orthocladius infuscatus Malloch, 1915b:
annulator. Ont., Calif., N. Mex.; PA. 517.-Ill.
Cricotopus annulator Goetghebuer, 1927:. Cricotopus aratus Roback, 1957a: 73.-Pa.
52.-Belgium. Cricotopus ceris Roback, 7957a: 72.-Pa.
Cricotopus irwini Sublette & Sublette, lestralis. Alas., N.W.T., Wash., Grld.; PA.
l97l: 97.-Calif. Trichocladius lestralis Edwards, I924b:
baptistensis. Ont. 172.-Spitsbergen.
Cricotopus baptistensis Boesel, 1983: luciae. Ont.
87.-Ont. Cricotopus luciae LeSage in LeSage &
beringensis. Alas. Harrison, 1980: 91.-Ont.
Cricotopus beringensis Oliver & Dillon, mackenziensis. N.W.T.
1988: 475.-Alas. Cricotopus mackenziensis Oliver, 1977 :
. Subgenus GYMNOMETRIOCNEMUS
Genus EURYHAPSIS Oliver Goetghebuer
Euryhapsis Oliver, 7981: 7ll.
Type-species, cilium Oliver (orig. des.). subnudus. B.C., Ont., N.S., Tenn., N.C.; PA.
Metiocnemus subnudus Edwards, 1929:
annuliventris. Calif. 316.-England.
Metriocnemeus annulivenrrls Malloch, Gymnometriocnemus marionensis Sather,
191-5a: 46.-Calif. 1969: 729.-8.C.
cilium. Y.T., N.W.T., Alta.; PA.
Euryhapsis cilium Oliver, 1981:
714.-N.W.T.
CHIRONOMIDAE DE I-A REGION NEARCNQUE 27
fusistylus. Alas., N.W.T., Alta., Mont., Colo., camptophleps Edwards, 1,929) (orig.
Grld. des.).
Orthocladius fusistylus Goetghebuer, 1933:
26.-Grld. ?boreoalpina. B.C., Alta.; PA.
Hydrobaenus fusistylus var. fusistylus Smittia boreoalpina Goetghebuer, 1944:
(Goetghebuer), Sather, 7976:- 94. 43.-Austria.
Hydrobaenus fusistylus var. octomerus ?camptophleps. B.C.; PA.
Srether, 7976: 99. Spaniotoma camptophleps Edwards, 1929:
hudsoni. S. Dak. 353.-Great Britain.
Hydrobaenus hudsoni Sether, 1976: halvorseni. N.W.T., N.B.; PA.
131.-S. Dak. Rheosmittia halvorseni Cranston & Sather.
johannseni. Man., Ont., S. Dak., Minn., Ohio, 1986: 45.-Norwav.
N.Y., Md., Ark., N.C., S.C.
Orthocladius johannseni Sublette, 1967b:
504 (n. nam. for Orthocladius Genus LIMNOPHYES Eaton
nivoriundus Johannsen, 1934: 348, Limnophyes Eaton, 1875: 60.
preocc. Fitch, 1847).-N.Y. Type-species, pusillus Eaton (mon.).
Orthocladius domus Sublette, 1966:
594.-Md. asquamatus. N.Y., Grld.; PA.
Trissocladius hamiltoni Sether, 1969: Limnophyes eltoni var. asquamatus
42.-Man. Andersen, 1937 : 7 Z.-Grld.
laticaudus. Alas. atomarius. N.Y., Grld.; PA.
Hydrobaenus laticaudus Sether, 1976: Chironomus atomarius Zetterstedt, 1850:
80.-Alas. 3522.-Sweden.
martini. N.W.T., N.B.; PA. borealis. Y.T. to Grld., N.8., s. to Minn.,
Hydrobaenus martini Sether, 1976: Tenn., N.C., S.C.; PA.
134.-N.W.T. Limnophyes borealis Goetghebuer, 1933:
pilipes. Ont., Qu., S. Dak., Ohio, Ill., Calif., 29.-Grld.
N. Mex., N.C., S.C., Fla. Limnophyes spatulosus Srether, t975d:
Orthocladius pilipes Malloch, 1915b: 1055.-Minn.
522.-\1. carolinensis. Ont., Minn., Tenn., N.C., S.C.
pilipodex. S. Dak., Ark., Ala. Limnophyes carolinensis Srether, I97 5d:
Hydrobaenus pilipodex Sether, 1976: 10-50.-s.c.
144.-5. Dak. coloradensis. Colo.
scapulapilosus. Alas., Ont. Limnophyes coloradensis Sether, 197 5d:
Hydrobaenus scapulapilosus Sether, L976: 1052.-Colo.
124.-Ont. eltoni. Grld.: PA.
spinnatis. Ont., Qu6.; PA. Camptocladius eltoni Edwards, L922:
Hydrobaenus spinnatis Sather, 1976: 203.-Bear Island.
152.-Qu. fumosus. Ont., N.S., N.Y., N.C., S.C., Ga.
travisi. Mass. Camptocladius fumosus Johannsen, 1905:
Hydrobaenus travisi Sa.ther, 1989b: 261.-N.Y.
55.-Mass. Limnophyes cristatissimus Sether, 7975d:
tumidistylus. N.W.T. 1032.-S.C.
Hydrobaenus tumidistylus Sether, 1976: globifer. Y.T., N.W.T., Alta., Man., Ont.,
160.-N.W.T. Grld.: PA.
virgo. Ont. Camptocladius globifer LundstrOm, 1915:
Hydrobaenus virgo Sather, L976: 16.-U.S.S.R.
128.-Ont. Limnophyes folliculatus Sether, L97 5d:
1050.-Man.
groenlandiensis. Grld.
Genus KRENOSMITTIA Thienemann & Limnophy e s gr o e nl and ie nsis Andersen,
Krtiger 70.-Grld.
1937.
Krenosmittia Thienemann & KrUger, L939: hamiltoni. Alas. to N.W.T., B.C., Ont., Calif.,
253. Type-species, Smittia gynocera Grld.
Edwards, 1937, sensu Thienemann & Limnophyes hamiltoni Saether, 1969:
Krriger (misident. = Spaniotoma 101.-B.C.
CHIRONOMIDAE DE LA REGION NEARCTIQUE 29
brevinervis. N.W.T.
Chironomus brevinervis Holmgren, Genus PARATRISSOCLADIUS Zavrel
1869: 39.-N.W.T. Paratissocladius Zavrel, 1937: 10 (as
Metriocnemus despectus Kieffer, 1926: subg. of Trissocladius Kieffer in
87.-N.W.T. Kieffer & Thienemann 1908).
debilipennis. N.Y., Grld. Type-species, Trissocladius fluviatilis
Chironomus debilipennis Lundbeck, 1898: Goetghebuer (orig. des.) =
286.-Grld. Chironomus excerptus Walker.
CHIRONOMIDAE DE LA REGION NEARCTIQUE -1-')
piloculus. N.W.T.
Genus TOKUNAGAIA Sether Hydrobaenus piloculus Cranston & Oliver,
Tokunagaia Sather, L973a: 56. 1988a: 435.-N.W.T.
Type-species, Spaniotoma kibunensis
Tokunaga (orig. des.).
Genus XYLOTOPUS Oliver
excellens. B.C.; PA. Xylotopus Oliver, 1982: 167. Type-species,
Eukiefferiella excellens Brundin, 1956: Orthocladius par Coquillett (orig. des.).
89.-Sweden.
obriaini. N.W.T. par. Ont. to N.S., s. to Iltah, Tex. & Fla.
Tokunagaia obriaini Hayes & Murray, Orthocladius par Coquillett, 1901b:
1988: 361.-N.W.T. 608.-N.J.
rowensis. Alta., Ont. Eurycnemus scitulus Coquillett, 1901b:
Adactylocladius rowensis Srther, 1969: 608.-N.J.
58.-Alta.
40 NEARCTIC CHIRONOMIDAE
lobus. Ky., N.C., s. to Miss. & Fla. pagana. B.C. to Qu., N.Y., Mich., S. Dak.
Chironomus lobus Beck, 7962: 89.-Fla. Miss.; PA.
lucifer. Sask., Ont., widespread U.S.A. Chironomus paganus Meigen, 1838:
Chironomus lucifer Johannsen, 1907b: 7.-Belgium.
110.-Kans. Einfeldia synchrona Oliver, 1971:
Chironomus indistinctus Malloch, 1915b: 1591.-Ont.
477.-I!]..
milleri. Man., Ont., Oreg., Minn., Mich., Ind.,
N.Y. Genus ENDOCHIRONOMUS Kieffer
Tendipes milleri Townes, 1945: 110.-N.Y. Endochironomus Kieffer, 1918b: 69.
modestus. Widespread NE., Grld.; PA. Type-species, Chironomus alismatis
Chironomus modestus Say, 1823: 13.-Pa. Kieffer (orig. des.) = Tipula tendens
neomodestus. N.W.T. to Qud., s. to Tex. & Fabricius.
Fla.
Chironomus neomodestus Malloch, 1915b: dispar. Not NE.
475.-\t. Chironomus dispar Meigen, 1830:
Chironomus tenellus authors, not 247.-Not stated.
Zetterstedt. oldenbergi. Grld.; PA.
nervosus. Alas., N.W.T., B.C. to N.B., s. to Endochironomus oldenbergi Goetghebuer,
Calif. & Fla., Grld.; PA. I932b: 130.-Europe.
Chironomus nervosus Staeger, 1839: nigricans. N.W.T., B.C. to N.B., s. to Calif.
567.-?Denmark. & Fla.
Chironomus brevitibialis Zetterstedt, 18-50: Chironomus nigricans Johannsen, 190-5:
3-535.-Sweden. 219.-N.Y., N.J.
simpsoni. N.Y., Ga., Fla. Chironomus johnsoni Kieffer, l9}6a: 19
Dicrotendipes simpsoni Epler, 1987: (n. nam. for Chironomus albipennis
84.-N.Y. Meigen of Johnson, 1900, nom. nud.).
thanatogratus. Fla. Chironomus albipennis authors, not
Dicrotendipes thanatogratrzs Epler, 1987: Meigen.
88.-Fla. subtendens. B.C. to Qu6. & Nfld., s. to N.
tritomus. Alas., Sask., Mont. to N.H., Okla., Mex. & Fla.
Ky., N.C., s. to La. & Fla.; pA. Tanytarsus subtendens Townes, 1.94-5:
Chironomus tritomus Kieffer in 65.-N.Y.
Thienemann & Kieffer, 1916: Chironomus viridis authors, not Macquart.
-524.-Sweden.
Chironomus incurvus Sublette,
1964b: 126.-La. Genus ENDOTRIBELOS Grodhaus
Endotribelos Grodhaus, 1987a: 237. Type-
species, Tendipes hesperius Sublette
Genus EINFELDIA Kieffer (orig. des.).
Einfeldia Kieffer, I924a: 393.
Type-species, pectoralis Kieffer (mon.). hesperium. Calif., Fla.
Tendipes hesperius Sublette, 1960:
austini. N. Mex. to N.C., s. to Fla. 217.-Calif.
Einfeldia austini Beck & Beck, 1970:
40.-Fta.
brunneipennis. Minn., Iowa to Mass., N.J., Genus GILLOTIA Kieffer
Ky., N.C., s. to Miss. & Fla. Gillotia Kieffer, 1927c: 272. Type-species,
Chir o nomus brunne ipe nnis Johannsen, Gillotia fuscipes Kieffer, 1922d (sub.
1905: 20-5.-N.Y. mon.; preocc. Kieffer, 1909) =
chelonia. N.Y., N.J., D.C., N. Mex. to La. Chironomus trifidus Freeman, 1957
Tendipes chelonia Townes, 194-5: (replacement nam.).
114.-N.Y.
natchitocheae. La. alboviridis. Iowa, Ill., Kans., N. Mex., Okla.
Chironomus natchitocheae Sublette, 1964b: Chironomus alboviridis Malloch. 191-5b:
124.-La. 482.-Tl/.
A1
CHIRONOMIDAE DE I-A REGION NEARCTIQUE
babiyi. Man. to Qu6., N.Y., Mich., Ohio, directus. Kans., Ky., N.C., s. to La. & Fla.
Okla., Tex., N.C., S.C., Fla. Tendipes directus Dendy & Sublette, 1959:
Chironomus babiyi Rempel, I937b: -514.-Ala.
274.-N.Y. elodeae. Qu6., N.Y.
bicorne. N.C., S.C., Fla. Harnischia elodeae Townes. 1945:
Kribioxenus bic ornis Townes, 194-5: 156.-N.Y.
35.-S.C. forceps. Man., Mich., Mass.
mirabile. Qud., N.Y., Fla. Harnischia forceps Townes, 194-5:
Krib ioxenus mirabilis Townes, 194-5: 157.-Mass.
35.-N.Y. frequens. Widespread NE.; PA.
Chironomus frequens Johannsen, 190-5:
230.-N.Y.
Genus OMISUS Townes hirtalatus. Ill. to Pa.. s. to Miss. & Fla.
Omisus Townes, 1945: 27. Type-species, Chironomus hirtalatus Beck & Beck, 1964:
pica Townes (orig. des.). 203.-Fla.
monochromus. Sask., Qu6., Mich., Ill. to
pica. Idaho, Man. to Qu6., N.H. to Conn., Mass., s. to Calif. & Fla.; PA.
Ky., N.C., s. to Miss. & Fla. Chironomus monochromus Wulp, 1874:
Omisus pica Townes, 1945:. 27.-N.H. 129 (n. nam. for Chironomus unicolor
Wulp, 1858: 162, preocc. Walker,
1848).-Netherlands.
Genus PAGASTIELLA Brundin pectinatellae. Ill. to Pa., s. to Miss. & Fla.
Pagastiella Brundin, 1949:. 840. Cryptochironomus pectinatellae Dendy &
Type-species, Chironomus orophilus Sublette, 19-59: -516.-Ala.
Edwards (orig. des.). potamogeti. Man. to Qu6., s. to N. Mex. &
Fla.
orophila. Maine to Conn., Ky., N.C., s. to Harnischia potamogeti Townes, 1945:
Miss. & Fla. 159.-N.Y.
Chironomus orophilus Edwards, schneideri. Ky., N.C., s. to Miss. & Fla.
1929: 405.-Great Britain. Parachironomus schneideri Beck & Beck,
ostansa. N.W.T., B.C. to Ont., N.H., N.C., 1969: 287.-Fla.
S.C. sublettei. Fla.
Paralauterborniella ostansa Webb, 1969: Chironomus sublettei Beck, 1961:
102.-Ont. 127.-Fla.
Paralauterborniella ostensa (error), tenuicaudatus. B.C., Sask. to Qu6.,
Boerger et al., 1982: 917.-Alta- widespread U.S.A.; PA.
Chirono mus tenuic aud atus Malloch.
1915b:475.-I].l.
Genus PARACHIRONOMUS Lenz varus. Man., N.Y.; PA.
Parachironomus LeDz, 1927: 160. Chironomus varus Goetghebuer, 1921:
Type-species, Chironomus cryptotomus 162.-Belgium.
Kieffer (Townes, L945: 152)-
galaptera. B.C., Man., Qu6., Ont., S. Dak. to duplicatus. Ont., S. Dak., Mich. to Okla.,
N.Y. & III. R.I., N.C., S.C.
Harnischia galaptera Townes, 1945: 148 Chironomus duplicatus Johannsen, I937b:
(n. nam. for Chironomus claripennis 28. (n. nam. for Chironomus basalis
Malloch, 191-5b: 439, preocc. Malloch, 1915b: 441, preocc. Staeger,
Lundbeck, 1898).-Mich. 1845).-N.Y.
loganae. Kans., Ohio, Ky., N.C., s. to Miss. &
Fla. fuscitibia. Calif.
Paracladopelma loganae Beck & Beck, Paratendipes fuscitibia Sublette, 1960:
1969: 303.-Fla. 200.-Calif.
nais. Ont., N.Y., Mich., Pa. nitidulus. Ohio, Mich., N.J., Pa., Del., Md.
Harnischia nais Townes, 1945: 149.-N.Y. Chironomus nitidulus Coquillett, 190lb:
S.C.,
nereis. Sask., Minn., Mich., Kans., N.C., 608.-N.J.
Ga.; PA. Chironomus nitidellus (error), Malloch,
Harnischia nereis Townes, 1945: 1915b: 468.
150.-S.C. nudisquama. Ont.; PA.
nixe. Colo. Chironomus nudisquama Edwards, 1929:
Harnischia nrxe 'fownes, \945: 149.-Colo. 396-England, Ireland.
undine. Alta. to N.Y., Oreg., S. Dak., Kans., subaequalis. Man. to Qu6., s. to Kans. &
Ky., N.C., s. to Miss. & Fla.; PA. Fla., Calif.; PA.
Harnischia undine Townes, 194-5: Chironomus subaequalis Malloch, 191-5b:
149.-Oreg. 440.- Ill.
winnelli. Y.T., N.W.T., Ont., Wis., Mich. thermophilus. Calif., Nev., Ariz., N. Mex.
Paracladopelma winnelli Jackson, 1977: Paratendipes thermophilus Townes, 1.945:
1334.-Mich. 31..-Ariz.
profusa. Wash. to Mont., s. to Calif. & N. aviceps. Alta., Ont., Qu6. to N.H., Mont. s.
Mex. to Calif. & Tex., Fla.
Tanytarsus profusus Townes, 1945: Polypedilum aviceps Townes, 1945:
73.-Nev. 61.-N.Y.
punctipes. Sask. to Ont., Qu6., N.Y., Idaho, braseniae. N.W.T. to Ont., N.Y., Ohio,
N.J., N.C., S.C.; PA. Wash., Oreg., S. Dak., Ky., Va. to Fla.
Chironomus punctipes Wiedemann, 1817: Chironomus bras eniae Leathers. L922:
65.-Germany. 8.-N.Y.
vittata. Ont., Qu6., N.J., N.C.,
S.C. californicum. Calif.
Tanytarsus vittatus Townes, 1945: Polypedilum californicum Sublette, 1960:
77.-N.J. 204.-Calif.
calopterus. Ont., Qu., Mich., N.Y., Md., Va.
Chironomus calopterus Mitchell, 1908:
Genus POLYPEDILUM Kieffer 8.-Md.
Polypedilum Kieffer, l9l2b: 4I. Polypedilum walleyi Townes, 1945:
Type-species, pelostolum Kieffer = 52.-Qu6.
Chironomus nubifer Skuse (Ashe, 1981: cinctum. Oreg., Calif., Nev., Va.
58). Polypedilum cinctum Townes, 1.94-5:
59.-Nev.
Subgenus PENTAPEDILUM Kieffer convictum. Sask., Ont., Qu6., s. to Kans. &
Pentapedilum Kieffer, l9I3b: 25. Fla.: PA.
Type-species, Pentapedilum stratiotale Chironomus convictus Walker, 1956a:
Kieffer (Edwards, 1929: 376) = 161.-England.
Chironomus tritus W alker. Chironomus flavus Johannsen, 1905:
225.-N.Y.
sordens. Qu6., Mich. to N.Y., R.I., Va.; PA. Chironomus flaviventris Johannsen, 1907b:
Tanytarsus sordens Wulp, 1874: 111.-Kans.
1.41.-Netherlands. Polypedilum obtusum Townes, 1945:
Tanytarsus fulvescens Johannsen, 1905: 60.-N.Y.
293.-N.Y. fallax. Alta., Sask., Qu6., s. to Utah & Fla.
Calopsecffa americ ana Kieffer, 1917: Chironomus fallax Johannsen, 190-5:
357.-N.Y. 210.-N.Y.
tritum. B.C. to N.8., s. to Calif., Tex. & Fla.; Polypedilum fallax var. fallax Johannsen,
PA. t905: 210.-Ont., Qu., Ill., Ind.,
Chironomus titus Walker, 1956a: Mich., Vt., N.Y., Tenn.
342.-England. fuscipenne. 8.C., Wash., Oreg., Mont. to
Polypedilum albulum Townes, 1945: Colo.; PA.
63.-N.Y. Chironomus fuscipennis Meigen, 1818:
35.-Europe.
Subgenus POLYPEDILUM Kieffer illinoense. N.W.T. to Qu6., s. to Calif. &
Fla.; PA.
albicorne. B.C., Ont., Qu., Nfld., Wis. to Chironomus illinoensis Malloch. 1915b:
Ohio & Mass., Idaho, Calif. to La., 471.-ilL
Fla. laetum. Alta. to Qu6., s. to Calif., N. Mex.,
Chironomus albicornis Meigen, 1838: N.C., S.C.; PA.
6.-Europe. Chironomus laetus Meigen, 1818:
Polypedilum nigritum Townes, 1,945: 38.-Europe.
56.-N.Y. nubeculosum. N.W.T. to Qu6., Minn. to
angustum. 8.C., Ont., Qu6., Nfld., Minn. to N.Y., Idaho, Ind., Colo.; PA.
Va., Calif., Ark., Fla. Chironomus nubeculosus Meigen, 1804:
Polypedilum angustum Townes, L94-5: 18.-Europe.
56.-N.J. ontario. Ont., Qu6. s. to N. Mex., La. &
artifer. Wash., Calif. to Qu6., N.Y., Ohio, N. D.C.
Mex.. Va. Chironomus ontario Walley, 1926b:
Chironomus artifer Cvrran, 1930: 206--Ont-
32.-N.Y. Chironomus hirtipes Mitchell, 1908: 9
(preocc. Macquart, 1834).-Md.
52 NEARCTIC CHIRONOMIDAE
Tribe TANYTARSINI
Tribe PSEUD O CHIRONOMINI
Genus CLADOTANYTARSUS Kieffer
Genus PSEUDOCHIRONOMUS Malloch Cladotanytarsas Kieffer, I92lc: 277.
Pseudochironomus Malloch, 191-5b: -500. Type-species, pallidus Kieffer (sub.
Type-species, richardsoni Malloch (orig. mon.; Kieffer, 1922e: 100).
des.). Lenziella Kieffer, 1922c: 36I.
Type-species, bicornuta Kieffer (orig.
anas. Qu6., N.Y. des.).
Pseudochironomus anas Townes. 7945:.
17.-N.Y. aeiparthenus. Ont.
articaudus. Man., Ont., Qu6. Cladotanytarsus aeiparthenus Bilyj,
Pseudochironomus articaudus Seether, 1989: 948.-Ont.
1977b: 81.-Man. atridorsum. Man.; PA.
aureus. Kans. Cladotanytarsus atridorsum Kieffer, 1924b:
Psectrocladius aureus Johannsen, 1908: 38.-Germany.
283.-Kans. crusculus. S. Dak., N. Mex.
badius. Man. Lenziella cruscula Sether, 1971: 1825.-5.
Pseudochironomus badius Sether 1977b: Dak.
69.-Man. daviesi. Ont.
banksi. Ont., N.J., Mass., Fla. Cladotanytarsus daviesi Bilyj' 1989:
Pseudochironomus banksi Townes, 1945: 9-51.-Ont.
17.-Mass. elaensis. Ont.
chen. Ont., Md., N. Mex. to La. Cladotanytarsus elaensis Bilyj, 1989:
Pseudochironomus chen Townes. 1945: 955.-Ont.
18.-Md. fusiformis. Ont.
crassus. N.W.T., Man. Cladotanytarsus fusiformis Bilyj,
Pseudochironomus crassus Townes, 1945: 1989: 9-54.-Ont.
15.-Hudson Bay Territory, N.W.T. mancus. Iowa: PA.
fulviventris. Man. to N.8., S. Dak. to N.Y., Chironomus mancus Walker, I956a:
s. to Tex. & Fla. 161.-England.
Chironomus fulviventris Johannsen, 1.905: muricatus. Ont.
229.-N.Y. Cladotanytarsus muricatus Bilyj'
julia. Qu., Mich., N.Y., Ohio, N. Mex. to 1989: 956.-Ont.
La. pinnaticornis. Ont.
Orthocladius julia Cwran, 1930: 34.-N.Y. Cladotanytarsus pinnaticornis Bilyj,
Pseudochironomus alx Townes. 1945: 1989: 953.-Ont.
19.-Mich.
56 NEARCTIC CHIRONOMIDAE
40.-Grld.
norvegicus. N.W.T.; PA.
Fournieria norvegica Kieffer, 1924a:
394.-Norway.
pallidicornis. N. Mex.; PA.
Chironomus pallidicornis Walker, 1956a:
169.-England.
quadratus. N. Mex. to N.C., s. to Fla.
Tanytarsus quadratus Sublette, 1,964b:
144.-La.
quadridentatus. Man.; PA.
Tanytarsus quadridentatus Brundin, 1947:
75.-Sweden.
recens. N. Mex. to N.C.. s. to Fla.
Tanytarsus recens Sublette, 1964b.
743.-La.
signatus. Ont., N.Y.; PA.
Chironomus signatus Wulp, 1858:
169.-Netherlands.
tibialis. Ont.
Tanytarsus tibialis Webb, 1969: 100.-Ont.
trilobus. Ont.
Tanytarsus tilobus Webb, 1969: 99.-Ont.
usmaensis. Man.; PA.
Tanytarsus usmansis Pagast, 193 1:
232.-U.S.S.R.
varelus. Pa.
Calopsectra varela Roback, 1957a:
128.-Pa.
Dendy, J.S., & J.E. Sublette. 19-59. The Fabricius,J.C. 178I. Species insectorum
Chironomidae (= Tendipedidae: Diptera) of exhibentes eorum differentias specificas,
Alabama with descriptions of six new species. synonyma, auctorum, loca natalia,
Ann. ent. Soc. Am. 52: 506-519. metamorphosin. 517 pp. Ifamburgi et
Eaton, A.E. 1875. Breves dipterarum uniusque Kilonii [=Hamburg and Kiel].
lepidopterarum Insulae Kerguelensi Fabricius, J.C. I794. Entomologia systematica
indigenarum diagnoses. Entomologist's mon. emendata et aucta. Secundum classes,
Mag. 12: 58-61. ordines, genera, species, adjectis synonymis,
Edwards, F.W. 1922. Results of the Oxford locis, descriptionibus, observationibus. 4,
University Expedition to Spitzbergen, 1921. 472 pp. Copenhagen.
Ann. Mag. nat. Hist. (9)lO: t93-2t5. Fabricius, J.C. 180-5. Systema antliatorum
Edwards, F.W. I924a. A note on the genus secundum ordines, genera, species. 373 pp-,
Protanypus Kieffer (Diptera: Chironomidae). + 30 pp. Brunsvigae [=Brunswick].
Annfs Biol. lacustre 13: 7I9-I22. Ferrington, L.C. & O.A. Sether. 1987. Male,
Edwards, F.W. I924b. Results of the Merron female, pupa and biology of. Oliveridia
College Expedition to Spitzbergen, 7923, hugginsi n. sp. (Chironomidae: Diptera) from
No. 4. Diptera Nematocera. Ann. Mag. nat. Kansas. J. Kans. ent. Soc. 6O: 457-461
Hist. (9)14: 162-174. Fitch, A. 7847. Winter insects of eastern New
Edwards, F.W. 1924c. Some British species of York. Am. J. Agric. Sci. 5: 274-284.
C orynoneura (Diptera-Chironomidae). Fittkau, E.J. 1957. Thienemannimyia und
Entomologist's mon. Mag. 60: 182-189. Conchapelopia, zstei neue Gattungen
Edwards, F.W. 7929. British non-biting midges innerhalb der Ablabesmyia-Costalis-Gruppe
(Diptera, Chironomidae). Trans. R. ent. (Diptera, Chironomidae).
Soc. Lond. 7'7: 279-430. Chironomidenstudien VII. Arch. Hydrobiol.
Edwards, F.W. 1931. Diptera of Patagonia and 53: 313-322.
South Chile. 11, 5, Chironomidae: 233-331.. Fittkau, E.J. 1962. Die Tanypodinae (Diptera:
Trustees of the British Museum, London. Chironomidae). Die Tribus Anatopyniini,
Edwards, F.W. 1932a. Notes on Highland Macropelopiini und Pentaneurini. Abh.
Diptera, with descriptions of six new species. Larvalsyst. Insekten 6: 7-453.
Scott. Nat. 794: 43-52. Fittkau,E.J. 7965. Revision der von E. Goeldi
Edwards, F.W. I932b. Recent literature, Faune aus dem Amazonasgebiet beschriebenen
de France: 23, Diptdres: Chironomidae, IV. Chironomiden (Diptera).
Par M. Goetghebuer. Paris (Lechavalier), Chironomidenstudien X. Beitr. neotrop.
1932. Entomologist 65: 140-141. Fauna 4: 209-226.
Edwards, F.W. 1933. Oxford University Fittkau, E.J. 1968a. Eine neue
Expedition to Hudson's Strait, 1931: Diptera Tanypodinae-Gattung, Dj almab atis ta
Nematocera. With notes on some other (Chironomidae, Diptera), aus dem
species of the genus Diamesa. Ann. Mag. brasilianischen Amazonasgebiet. Amazoniana
nat. Hist. 10(12): 6Il-620. l:327-349.
Edwards, F.W. 1935. Diptera Nematocera from Fittkau,E.J. 1968b. Siolimyia amazonica n.
East Greenland. Ann. Mag. nat. Hist. gen. n. spec., eine flugfdhige Chironomide
10(15):467-473. (Diptera) mit linem Hypopygium inversum.
Edwards, F.W. 1937. Chironomidae (Diptera) Amazoniana L: 259-265.
collected by Prof. Aug. Thienemann in Fittkau, E.J. & D.A. Murray. 198-5.
Swedish Lappland. Ann. Mag. nat. Hist. Ser. Radotanypus a new genus of Tanypodinae
10(20): 140-148. from the Nearctic (Diptera, Chironomidae).
Edwards, F.W. 1938. I. A new genus of the Spixiana Suppl. (Muench.) 17: 209-273.
subfamily Podonominae. Pp. 152-154. In: Fittkau, E.J. & D.A. Murray. 1988.
Edwards, F.W. & A. Thienemann. Neuer Bethbilbeckia floridensis: a new genus and
Beitrag zur Kenntnis der Podonominae species of Macropelopiini from the South
(Dipt. Chironomidae). Chironomidae aus Eastern Nearctic (Diptera: Chironomidae).
Lappland IV. Zool. Anz. 122: 752-758. Spixiana Suppl. (Muench.) 74: 253-259.
Enderlein, G. 1972. Die Insekten des Antarkto- Fleutiaux, E. 1933. Genre nouveau et especes
Archiplata-Gebietes. K. svenska nouvelles d'Eucnemididae de Madagascar
VetenskAkad. Handl. 48: 106-110. (Col.). Bull. Soc. ent. Fr. (1932): 286-288.
Epler, J.H. 1987. Revision of the Nearctic Freeman, P. 1957. A study of the
Dicrotendipes Kieffer, 1913 (Diptera: Chironomidae (Diptera) of Africa south of
Chironomidae). Evol. Monogr. 9: i.-1.02.
CFIIRONOMIDAE DE LA REGION NEARCTIQUE
the Sahara. Part III. Bull. Br. Mus. nat. Goetghebuer, M. 195-5. Deux Chironomides
Hist. (Ent.) 5: 321-426. nouveaux du Groenland (Dipt. Nemat.).
Fries, B.F. 1830. Beskrifning 6fver ett nytt Expeditions polaires francaises (Groenland):
sllgte Hydrobaenus hOrande till Tipulariae. (Collection A. Bauer du Musee Zoologique
K. svenska VetenskAkad. Handl. 1829: de Strasbourg). Bull. Ass. philomath. Als.
176-187. Lorr. 9: 767.
Garrett, C.B.D. 1925- Seventy new Diptera. Goetghebuer, M.& F. Lenz. 7936.
Key to Pseudoleria, Helomyzidae, Tipulidae, Tendipedidae-Pelopiinae (Chironomidae-
Chironomidae, Dixinae, Orphnephilidae, Tanypodinae). Fliegen palaearkt. Reg. 13b:
Mycetophilidae. Cranbrook, B.C., 16 pp. 1-81.
Gistl, J. 1835. Neue Arten von Dipteren aus Goetghebuer, M.& F.Lenz. 1937.
der Umgegend von Mrinchen, benannt und Tendipedidae-Tendipedidae
beschrieben von Meigen, aufgefunden von (Chironomidae-Chironominae). [part].
Dr. J. Waltl. Faunus 2: 66-72. Fliegen palaearkt. Reg. 13c: 1-48.
Goeldi, E.A. 190-5. Os mosquitos no Para. Goetghebuer, M. & F. Lenz- 1938.
Mems Mus. paraense Hist. nat. Ethnogr. 4: Tendipedidae-Tendipedidae
734-r39. (Chironomidae-Chironominae). [part].
Goetghebuer, M. 1921. Chironomides de Fliegen palaearkt. Reg. 13c: 73-128.
Belgique et sp6cialement de la zone des Goetghebuer, M. & F. Lenz. 1939a.
Flandres. M6m. Mus. r. Hist. nat. Belg. 8: Tendipedidae-Diamcsinae
r-211. (Chironomidae-Diamesinae). Fliegen
Goetghebuer, M. 1922. Nouveaux matdriaux palaearkt. Reg. 13d: 1-30.
pour l'6tude de la fauna des Chironomides Goetghebuer, M. & F. Lenz. 1939b.
de Belgique. Annls Biol. lacustre ll:. 38-62. Tendipedidae-Corynone urinae
Goetghebuer, M. 1927 - Les Cricotopus de (Chironomidae-Corynoneurinae). Fliegen
Belgique (Dipt. Chironomides). Bull. Annls palaearkt. Reg. 13f: 1-39.
Soc. r. ent. Belg. 67: 5l-54. Goetghebuer, M. & F. Lenz. 1940.
Goetghebuer, M. 1928. Diptdres Tendipedidae-Orthocladiinae
(Ndmatocdres): Chironomidae. IIL (Chironomidae-Orthocladiinae) [part].
Chironomariae. Faune Fr. 18: 1-174. Fliegen palaearkt. Reg. 13g: 1-24.
Goetghebuer, M. 1931. Ceratopogonidae et Goetghebuer, M. & F- Lenz. 1943.
Chironomidae nouveaux d'Europe. Bull. Tendipedidae-Orthocladiinae
Annls Soc. r. ent. Belg. Tl; 217-218. (Chironomidae-Orthocladiinae) [part].
Goetghebuer, M. 1.932a. Diptdres Fliegen palaearkt. Reg. 13g: 65-t1,2-
Chironomidae 1V (Orthocladiinae, Goetghebuer, M. & F. Lenz. 1,944.
Corynoneurinae, Clunioninae, Diamesinae). Tendipedidae-Orthocladiinae
Faune Fr.23: 1.-204. (Chironomidae-Orthocladiinae) fpart].
Goetghebuer, M. 7932b. Ceratopogonidae et Fliegen palaearkt. Reg. 13g: 113-144.
Chironomidae nouveaux ou peu connus Goetghebuer, M. & F. Lenz. 1954.
d'Europe. Bull. Annls Soc. r. ent. Belg. 72: Tendipedidae-Tendipedidae
t25-r30. (Chironomidae-Chironominae). [part].
Goetghebuer, M. 1933. Chironomides du Fliegen palaearkt. Reg. 13c: 128-168.
Groenland oriental, du Svalbard et de la Goetghebuer, M. & C.H. Lindroth. 1931. Fam.
terre de Frangois Joseph. Skr. Svalbard Ceratopogonidae und Chironomidae. Pp.
Ishavet 53: 19-31. 274-285. In: Lindroth, C.H. (Ed.). Die
Goetghebuer, M. 193-5. Ceratopogonidae et Insektenfauna Islands und ihre Problem.
Chironomidae rdcoltds pour la premidre fois Zool. Bidr. Upps. 13: 105--599.
en Belgique. Bull. Annls. Soc. r. ent. Belg. Gouin, F. t936. M6tamorphoses de quelques
75: 413-418. Chironomides d'Alsace et de Lorraine avec
Goetghebuer, M. 1936. Chironomides du Congo la description de trois espdces nouvelles par
Belge. Revue Zool. Bot. atu.28 453-492- M. Goetghebuer. Revue fr. Ent. 3: l5l-173-
Goetghebuer, M. L940. Chironomides de Gould, J. 1840. Birds of Australia, 1-80.
Laponie Su6doise. Bull. Annls Soc. r. ent. (Aves).
Belg. 80: 55-72. Gowin, F. 1943. Orthocladiinen aus Lunzer
Goetghebuer, M. 1944. Ceratopogonidae et Fliessengewdssern. II. Arch. Hydrobiol. 40:
Chironomidae nouveaux ou peu connus 114-122.
d'Europe (douziOme note). Biol. Jaarb. 11:
35-44.
NEARCTIC CHIRONOMIDAE
Mem. Cornell Univ. agric. Exp. Stn 205: nouveau. Bull. Soc. Hist. nat. Metz 27:
1-84. 103- 105.
Johannsen, O.A. t937b. Aquatic Diptera. Part Kieffer,J.J. 191,1c. Nouveaux Tendipddides du
IV. Chironomidae: Subfamily Chironominae. groupe Orthocladius (Dipt.). (1'note); (2*
Mem. Cornell Univ. agric. Exp. Stn 210: 3- note). Bull. Soc. ent. Fr. 1911: 181-187;
56. r99-202.
Johannsen. O.A. 1942. Immature and adult Kieffer,J.J. 19t2a. Nouveaux Chironomides
stages of new species of Chironomidae (Tendipedidae) de Ceylan. Spolia zeylan. 8:
(Diptera). Ent. News 53: 7O-75- l-24.
Johannsen. O.A. 7943. Adult and immature Kieffer, J.J. I9l2b. Tendipedidae (Dipt.). In:
stages of Cricotopus elegans n. sp. M. Sauter's Formosa Ausbeute. Supplta.
(Chironomidae, Diptera). Ent. News 54: ent. l.: 27-43.
77-79. Kieffer, J.J. 191.2c. Quelques nouveaux
Johannsen, O.A. 7946. Revision of the North Tendip6dides (Dipt.) obtenus d'dclosion. I.
American species of the genus Pentaneura note. Bull. Soc. ent. Fr. 17: 86-88.
(Tendipedidae: Chironomidae, Diptera). Jl Kieffer. J.J. l9l3a. Nouvelle contribution a la
N.Y. ent. Soc. 54: 267-289. connaissance des Tendip6dides D'Allemagne.
Johannsen, O.A. 1947. A new species of Bull. Soc. Hist. nat. Metz. 3: 37-44.
Hydrob aenus (Chaetocladius) fr om Kieffer, J.J. 1913b. Nouveaux Chironomides
Connecticut with notes on related forms (Tendip6dides) d'Allemagne. Bull. Soc. Hist.
(Diptera, Chironomidae). Ent. News 58: nat. Metz 28:. 7-35.
I7l-174. Kieffer, J.J. 1913c. Nouvelle 6tude sur les
Johnson, C.W. 1900. Order Diptera. Pp. 617- Chironomides de l'Indian Museum de
'700. In: Smith, J.B. Insects of New Jersey. Calcutta. Rec. Indian Mus. 9: ll9-I97.
A list of the species occurring in New Jersey Kieffer. J.J. 1913d. Chironomidae et
with notes on those of economic importance. Cecidomyidae. Result. scient. Voy. Ch.
Bull. N.J. agric. Ann. Rept. (1889) 27 Alluaud et R. Jeanell en Afrique Orientale
(Supp.): 1-75-5. (1e11-1012) (Dipt.) I: 1-43.
Johnson, C.W. 1925. Fauna of New England. Kieffer, J.J. 1913e. Dasyhelea halophila n. sp.,
15. List of the Diptera or two-winged flies. eine neue halophile Zuckmticke. Pp. 255-
Occ. Pap. Boston Soc. nat. Hist. 7: l-326. 256. In: Kieffer, J.J., & C. van Douwe
Jones, F.M. 1916. Two insect associates of the (Eds). Zur Flora und Fauna der
California pitcher plant, Darlingtonia Strandtrimpel von Rovigno (in Istrien). Biol.
californica (Dipt.). Ent. News 27: 385-392. Zbl. 33: 254-258.
Karunakaran, L. 1969. A new genus of the Kieffer. J.J. 1914. South African
subfamily Tanypodinae (Diptera, Chironomidae (Diptera). Ann. S. Afr. Mus.
Nematocera: Chironomidae) from Singapore. l0: 259-270.
Proc. R. ent. Soc. Lond. 38: 75-79. Kieffer, J.J. 1915a. Neue Chironomiden aus
Kellogg, V.L. 1900. An extraordinary new Mitteleuropa. Boteria Ser. zool. 13: 6-5-87.
maritime fly. Biol. Bull. 1: 81-87. Kieffer. J.J. 1915b. Fauna faeroensis: 2.
Kevan, D.K.McE. & F.E.A. Cutten-Ali-Khan. Tendipedidae (Chironomidae). Zool. Jb. 39:.
1975. Canadian Nymphomyiidae (Diptera). 103-720.
Can. J. Zool. 53: 8-53-866. Kieffer, J.J. 1917. Chironomides d'Amdrique
Kieffer, J.J. 1906a. Diptera Fam. conservds au Mus6e National Hongrois de
Chironomidae. Genera Insect. 42: I-78. Budapest. Budapest Magyar Nemzeti Muz.,
Kieffer, J.J. 1906b. Description d'un genre Ann. Hist. Nat. 15: 292-364.
nouveau et de quelques espBces nouvelles Kieffer,J.J. 1918a. Beschreibung neuer, auf
de Dipt0res de I'Amdrique du Sud. Annls Lazarettschiffen des 0stlichen
Soc. scient. Brux. 30: 349-358. Kreigsschauplatzes und bei Ignalino in
Kieffer, J.J. 1909. Diagnoses de nouveaux Litauen von Dr. W. Horn gessammelter
Chironomides d'Allemagne. Bull. Soc. Hist. Chironomiden mit Uebersichtstabellen einer
nat. Metz 26:. 37-56- Gruppen von paldarktischen Arten (Dipt.).
Kieffer, J.J. 1911a. Nouvelles descriptions de Ent. Mitt. 7: 35-53;94-IIO; 163-l7O; 177-
Chironomides obtenus d'dclosion. Bull. Soc. 188.
Hist. nat. Metz 27: l-60. Kieffer, J.J. 1918b. Chironomides d'Afrique et
Kieffer, J.J. 1911b. Description d'un d'Asie conservds au Mus6e National
Chironomide d'Amrique format un genre Hongrois de Budapest. Annls hist.-nat. Mus.
natn. hung. 16: 31-139.
66 NEARCTIC CHIRONOMIDAE
Kieffer, J.J. 1,919a. Chironomides d'Europe l'Europe. Annls Soc. scient. Brux. 43: 390-
conserv6s au Mus6e National Hongrois de 397.
Budapest. Budapest Magyar Nemzeti Muz., Kieffer, J.J. 1924b. Chironomides nouveaux ou
Ann. Hist. Nat. 17: 1-160. rares de l'Europe centrale. Bull. Soc. Hist.
Kieffer, J.J. 1919b. Observations sur les nat. Moselle 30 [-ser. 3,6]: 11-110.
Chironomides (Dipt.) ddcrits par J.R. Kieffer, J.J. 1924c. Quelques nouveaux
Malloch. Bull. Soc. ent. Fr. 7979: l9l-794. Chironomides de Scandinavie. Annls Soc.
Kieffer, J.J. 1,920a. Tableau synoptique des scient. Brux. 44: 80-86.
Chironomides pal6arctiques appartenant aux Kieffer, J.J. 1,925a- Deux genres nouveaux et
genres Polypedilum et Limnochironomus. plusieurs espCces nouvelles du groupe des
Annls Soc. scient. Brux. 39: 759-767. Orthocladiariae (Dipteres, Chironomidae).
Kieffer, J.J. I920b. IJn nouveau genre de Annls Soc. scient. Brux. 44: 55-5-566.
Chironomide (Dipt.). Bull. Soc. ent. Fr. Kieffer, J.J. 7925b. Nouveaux reprdsentants du
I9l9: 333-334. groupe Tanytarsus (Diptdres Chironomides)
Kieffer, J.J. 1921,a. Chironomides nouveaux ou de I'Europe centrale. Annls Soc. scient.
peu connus de la rdgion paldarctique. Bull. Brux. 44: 278-227.
Soc. Hist. nat. Moselle 29 [-ser.3,-5]: -51-109. Kieffer, J.J. 1.925c. Description de deux
Kieffer, J.J. 1921b. Chironomides de l'Afriqu6 Chironomides remarquables d'Estonie
6quatoriale (1'" partie). Annls Soc. ent. Fr. (Dipt.). Bull. Soc. ent. Fr. 30: 15-16.
90:1-56 Kieffer, J.J. 1926. Chironomiden der 2.
Kieffer, J.J. 1921,c. Synopse de la tribu des Fram-Expedition (1898-1902). Norsk ent.
Chironomariae (Diptdres). Annls Soc. scient. Tidsskr. (1925) 2: 78-89.
Brux. 40: 269-277. Kieffer, J.J. & W. Lundbeck. 1911. Diptera.
Kieffer, J.J. I922a. Diagnoses de nouveaux Pp. 272-275. In: Koenig, A. Avifauna
genres et espCces de Chironomides (Dipt.). spitzbergensis. Bonn. 294 pp.
Bull. Soc. ent. Fr. 7921,: 287-289. Kieffer. J.J. & A. Thienemann. 1906. Uber die
Kieffer, J.J. 1922b. Cbironomides de la Chironomidengattung Orthocladius. Z. wiss.
Nouvelle-Zemble. Rep. scient. Results InsektBiol. 2: L43-156.
Norw. Exped. to Novaya Zemlya 1927. 2: 1,- Kieffer, J.J. & A. Thienemann. 1908. I. Neue
24. und bekannte Chironomiden. Pp. 1-10, 33-
Kieffer, J.J. 1922c. Nouveau Chironomides a 39, 78-84. In: Kieffer, J.J. & A.
larves aquatiques. Annls Soc. scient. Brux. Thienemann. Neue und bekannte
4r-. 355-367. Chironomiden und ihre Metamorphose. Z.
Kieffer, J.J. I922d. Chironomides de l'Afrique wiss. InsektBiol. 4: 1-10, 33-39,78-84, 724-
Equatoriale (2' partie). Annls Soc. ent. Fr. I28, 184-190, 21,4-219, 256-259, 277 -286.
91: l-72. Kieffer, J.J. & A. Thienemann. I9I9.
Kieffer, J.J. 1922e. Chironomides nouveaux ou Chironomiden gesammelt von Dr. A. Koch
peu connus de la rdgion paldartique. Annls (Munster i.W.) auf den Lofoten, der
Soc. scient. Brux. 42: 71,-128. Bareninsel und Spitzbergen (Dipt.).
Kieffer, J.J. 1922t. Notice sur quelque ((Vorwort von A. Kock: 38-39)).
Chironomides d'Amdrique et de Nouvelle- Chironomiden der nordlichen Polarregion
Z6.lande. Annls Soc. linn. Lyon (1921) 68: (Kieffer, J.J.: 40-48; 110-120).
145-r48. Chironomidenlarven und -puppen von
Kieffer, J.J. 1923a- Diagnose de quelque Bareninsel un Spitzbergen. (Thienemann, A.:
nouveaux Tanypodines (Dipt.). Bull. Soc. 120-124). Ent. Mitt. 8: 38-48; 710-724.
ent. Fr. 296-297. Kruseman, G. 1933. Tendipedidae
Kieffer, J.J. 1923b. Chironomides de l'Afrique Neerlandicae. Pars I. Genus Tendipes cum
E,quatorial (3'partie). Annls Soc. ent. Fr. generibus finitimis. Tijdschr. Ent- 76: ll9-
92: 149-204. 216.
Kieffer, J.J. 7923c. Nouvelle contribution A Langton, P.H., P.S. Cranston & P. Armitage.
l'6tude des Chironomides de la 1988. The parthenogenetic midge of water
Nouvelle-Zemble. Rept. scient. Results supply systems, Paratanytarsus grimmii
Norw. Exped. Nova Zemlya 9: 3-11. (Schneider) (Diptera: Chironomidae). Bull.
Kieffer, J.J. I923d. Chironomides nouveaux ou ent. Res. 78:. 3I7-328.
peu connus de la rgion paldarctique. Annls Latreille, P.A. 1810. Considdrations gdndrales
Soc. scient. Brux. 42: 138-180. sur l'ordre naturel des animaux composant
Kieffer, J.J. 1924a. Quelques Chironomides les classes des Crustaces, des Arachnides et
nouveaux et remarquables du Nord de des Insectes; avec un tableau m6thodique de
CHIRONOMIDAE DE LA REGION NEARCTIQUE 67
leurs genres, dispos6s en familles. Paris, 444 Loew, II. 1866. Diptera Americae
pp- septentrionalis indigena. Centuria septima.
Leathers, A.L. 1922. Chironomus braseniae Berl. ent. Z. 10:. l-54.
new species (Dip., Chironomidae). Ent. Lundbeck, W. 1898. Diptera groenlandica.
News 33: 8. Naturhist. for. Kj@benhavn, Vidensk. meddel.
Lenz, F. L921. Chironomidenpuppen und lO: 236-314.
-larven. Bestimmungstabellen. Dt. ent. Z. LundstrOm, C. 1910. Beitrdge zur Kenntnis der
l92l: 148-162. Dipteren Finlands VI. Chironomidae. Acta
Lenz, F. l94la. Die Metamorphose der Soc. Fauna Flora fenn. 33:. 7-47.
Chironomidengattung Cryptochironomus. Lundstrdm, C. 1915. Resultats scientifiques de
Zool. Anz.133:.29-41. I'Expddition Polaire Russe en 1900-1903,
Lenz, F. 1941b. Die Jugenstadien der Sectio sous la direction du Baron E. Toll. Sect. E:
Chironomariae (Tendipedini) connectentes Zoologie 2, 8- Diptera Nematocera aus den
(Subf. Chironominae = Tendipedidae). arctischen Gegenden Sibiriens. [Akad. Nauk
Zusammenfassung und Revision. Arch. S.S.S.R.l Imp. Akad. Nauk Petrograd, Zap.,
Hydrobiol. 38: 1-69. Fiz.-Mat. Otd. (Acad. Imp. des Sci. de
Leonard, M.D. 1928. List of the insects of New Petsrograd, Me'm. Cl. Phys.-Math.) ser. 8,
York with a list of the spiders and cerain 29: l-33.
other allied groups. Mem. Cornell agric. Macquart, J. 1826. Insectes Diptdrds du Nord
Exp. Stn. (1926) 101:1-1121. de la France. I. Tipulaires. Soc. d'Amateurs
LeSage, L. & A.D. Harrison. 1980. Taxonomy Sci., Agr. Arts, Lille, Recueil Trav.
of Cricotopus species (Diptera: 182311824: 59-224.
Chironomidae) from Salem Creek, Ontario. Macquart, J. 1834. Historie Naturelle des
Proc. ent. Soc. Ont. lI7:. 57-714. Insectes. tome 1. 2. Diptdres. 578 pp.
Lindeberg, B. 1963. Taxonomy, biology and Macquart, J. 185-5. Diptdres exotiques
biometry of Tanytarsus curticornis Kieff. and nouveaux ou peu connus. Libr. Encycl.
T- brundini n. sp. (Dipt. Chironomidae). Roret. Suppl. 5: 35-1-56.
Suom. hydnt. Aikak. 29: 118-1.30. Makarchenko, E.A. 1982. Khironomidy roda
Lindeberg, B. 1967- Sibling species delimitation Protanypus Kieffer (Diptera, Chironomidae)
in the Tanytarsus lestagei aggregate (Diptera, dal'nego Vostoka SSSR. Pp. 124-744. In:
Chironomidae). Annls zool. Fenn. 4: 45-86. Biologiya presnovodnykh Zibotnykh dal'nego
Lindeberg, B. 1970. Tanytarsini ((Diptera, vostoka. Akad. NAUK. SSSR. biol. pochv.
Chironomidae) from northern Fennoscandia. Inst., Vladivostok.
Annls zool. Fenn. 7: 303-312- Makarchenko, E.A. 1983. Arctodiamesa gen.
Linnaeus, C. 1758. Systema naturae per regna nov. - Novyi rod khironomid podsemeistva
tria naturae, secundum classes, ordines, Diamesinae (Diptera, Chironomidae). In:
genera, species, cum caracteribus, Vsesoyuzneji simposium, Biologicheskie
differentiis, synonymis, locis. Ed. 1.0. 1., 824 problemy severa, Magadan 2: 264-265 .
pp' Holmiae (=Stockholm). Makarchenko, E.A. 1,986. Diamesa subletti sp.
Linnaeus, C. I767- Systema naturae, per regna n., a new species of chironomid (Diptera,
tria naturae, secundum classes, ordines, Chironomidae) from Canada. Aquat. Insects
genera, species, cum caracteribus, 8:155-1-57.
differentiis, synonymis, locis. Ed. 13, Malloch,J.R. 1915a. Four new North America
revised. 1(2): -533-1327- Holmiae Diptera. Proc. biol. Soc. Wash. 28; 45-48.
( =Stockholm). Malloch, J.R. 1915b. The Chironomidae, or
Lipina, N.N. 1939. Novye formy molodykh midges, of Illinois, with particular reference
stadii Chironomidae podopitnykh ozer to the species occurring in the Illinois River.
sapropelovoi stantsii v Zaluchie. Trudy Lab. Bull. Ill. St. Lab. nat. Hist. L0: 275-543.
sapropel. Otlozh. 1: 89-107. Malloch, J.R. 1918. A new species of
Loew, H. 1861a. Diptera. Americae Orthocladius (Chironomidae, Diptera). Bull.
septentrionalis indigena. Centuria prima. Brooklyn ent. Soc. 13:. 42.
Berl. ent. Z- 5: 307-359. Malloch, J.R. 1919. Part C. Diptera (excluding
Loew, tI. 1861b. Diptera aliquot in insula the Tipulidae and Culicidae). The Diptera
Cuba collecta. Wien. ent. Mschr. 5: 33-43. collected by the Canadian Arctic Expedition,
Loew, II. 1864. Diptera Americae 1913-1918. Rep. Can. Arctic Expedition
septentrionalis indigena. Centuria quinta. 1913-1918. Vol. III: Insects. pp. 34-90
Berl. ent. Z. 8: 49-104. Malloch, J.R. 1923. Insects, arachnids, and
chitopods of the Pribilof Islands, Alaska.
NEARCTIC CHIRONOMIDAE
Diptera (except Tipulidae, Rhyphidae and Chironomidae) from Japan. Konty0, Tokyo
Calliphoridae). N. Am. Fauna 46: 170-227. 55: 459-476.
Malloch, J.R. 19-14. The exploration of Nowicki, M.S. 1873. Beitrlige zur Kenntniss der
Southampton Island, Hudson Bay, by George Dipterenfauna Galiziens. 35 pp. Krakau,
Miksch Sutton, sponsored by Mr. John Poland.
Bonner Semple 1929-1930. Part II. de. 1967. Novo genero de
Oliveira, S.J.
Zoologie. Section 4. Spiders and insects (in Chironomidae da Amazdnia (Insecta,
part) of Southampton Island. III. Diptera). Atas Simp. Biota amazon. 5
Chironomidae, Sciaridae, Phoridae, (Zool.): 4r7 -479.
Syrphidae, Piophilidae, Helomyzidae, Oliver, D.R. 1959. Some Diamesini
Calliphoridae, Oestridae, and Tachinidae. 1. (Chironornidae) from the Nearctic and
Mem. Carneg. Mus. L2, pt. II, sect. 4: 1,3-32. Palaearctic. Ent. Tidskr. 90:. 48-64.
Mason, P.G. 1986. Four new species of the Oliver, D.R. 1971,. Description of Einfeldia
Cryptochironomus fulvus (Johannsen) species synchrona n. sp. (Diptera: Chironomidae).
complex (Diptera: Chironomidae). Can. Ent. 103: 1591-1595.
Entomologica scand. L6: 399-473. Oliver,D.R. 1976. Chironomidae (Diptera) of
Meigen, J.W. 1800. Nouvelle classification des Char Lake, Cornwallis Island, N.W.T., with
mouches A deux ailes (Diptera L.) d'apres un descriptions of two new species. Can. Ent.
plan tout nouveau. Paris, 40 pp. 108:10-53-1064.
Meigen, J.W. 1803. Versuch einer neuen Oliver, D.R. 1.977. Bicinclus-group of the genus
Gattungseinteitung der europAischen Cricotopus van der Wulp (Diptera:
zweiflrigetigen Insekten. Magazin Insektenk. Chironomidae) in the Nearctic with a
(Illiger) 2: 259-287. description of a new species. J. Fish. Res.
Meigen, J.W. 1804. Klassifikazion und Bd Can. 34:.98-104.
Beschreibung der europdischen zweiflrigeligen Oliver, D.R. 1981. Description of Euryhapsis
Insecten (Diptera Linn.). Erster Band. Abt. new genus including three new species
I., pp. I-I52. Abt. II, pp. 153-314. (Diptera: Chironomidae). Can. Ent. 113:
Braunschweig. 7lt-122.
Meigen, J.W. 1818. Systematische Beschreibung Oliver, D.R. 7982. Xylotopus, a new genus of
der bekannten europ5ischen, zweiflUgeligen Orthocladiinae (Diptera: Chironomidae).
Insekten. 1, xxxvi + 332 pp. Forstmann, Can. Ent. L1.4: 167-1,68.
Aachen. Oliver, D.R. 1.984. Description of a new species
Meigen, J.W. 1830. Systematische Beschreibung of Cricotopus van der Wulp (Diptera:
der bekannten europdischen zweiflrigeligen Chironomidae) associated with Myriophyllum
Insekten. 6: 1-401 pp. Schulz, Hamm. spicatum. Can. Ent. 116:. 1287-1292.
Meigen, J.W. 1838. Systematische Beschreibung Oliver, D.R. & M.E. Dillon. 1988. Review of
der bekannten europtiischen zweiflrigeligen Cricotopus (Diptera: Chironomidae) of the
Inseckten. 7: 7-434. Schulz, Hamm. Nearctic zone with descriptions of two new
Meunier, F. 1904. Monographie des species. Can. Ent. 120. 464-496.
Cecidomyidae, des Sciaridae, des Oliver. D.R. & M.E. Roussel. 1983.
Mycetophilidae et des Chironomidae de Redescription of Brillia Kieffer (Diptera:
I'ambre de la Baltique. Annls. Soc. scient. Chironomidae) with descriptions of Nearctic
Brux. 28: 1,2-274. species. Can. Ent. Il5:. 257-279.
Michailova. P. & J. Fischer 1986. Chironomus Osten Sacken, C.R. 7877. Western Diptera:
vancouveri sp. n. from Canada (Diptera, Descriptions of new genera and species of
Chironomidae). Reichenbachia 23: 99-106. Diptera from the region west of the
Mitchell, E.G. 1908. Descriptions of nine new Mississippi and especially from California.
species of gnats. Jl N.Y. ent. Soc. t6: 7-I4. Bull. U.S. geol. geogr. Surv. Terr. 3:
Murray, D.A. & E.J. Fitrkau. 1985. Hayesomyia 189-354.
a new genus of Tanypodinae from the Packard, A.S. 1869. On insects inhabiting salt
Holarctic (Diptera: Chironomidae). Spixiana water. Proc. Essex Inst. 6: 41-51
Suppl. (Muench.) lI: 195-207. Pagast, F. 1931. Chironomiden aus der
Muttkowski. R.A. 191-5. New insect life Bodenfauna des Usma-Sees in Kurland.
histories. L Bull. Wis. nat. Hist. Soc. 13: Fofia zool. hydrobiol. 3: L99-248.
L09-r22. Pagast, F. 1947. Systematik und Verbreitung
Nishida, H. 1987. Pseudocricotopus, a new der um die Gattung Diamesa gruppierten
subgenus of the genus Cricotopus (Diptera,
CHIRONOMIDAE DE LA REGION NEARCTIQI]E
Sather, O.A. 1975b. Two new species of (Diptera: Chironomidae) from South
Heterotanytarsas Spflrck, with keys to Carolina. Aquat. Insects 3: 193-198.
Nearctic and Palaearctic males and pupae of S
the genus (Diptera: Chironomidae). J. Fish. Chironomidae) from SE U.S.A., with
Res. Bd Can. 32: 259-270. descriptions of Plhudsonia, Unniella and
Sether, O.A. 1975c. Two new species of Platysmittia n. genera and Atelopodella n.
Protanypus Kieffer, with keys to Nearctic and subgen. Entomologica scand. 13: 46-5-510.
Palaearctic species of the genus (Diptera: Seether, O.A. 1983a. Oschia dorsenna n. gen.,
Chironomidae). J. Fish. Res. Bd Can- 32: n. sp. and Saetheria hirta n.sp., two new"
367-388. members of the Harnischia comPlex
Sether, O.A. 1975d. Twelve new species of (Diptera: Chironomidae). Entomologica
Limnophyes Eaton, with keys to Nearctic scand. 14: 395-404.
males of the genus (Diptera: Chironomidae). Srther, O.A. 1983b. A review of Holarctic
Can. Ent. 107: 1.029-1056. Gymnometriocnemus Goetghebuer, 1932, with
Sather, O.A. I976. Revision of Hydrobaenus, the description of Raphidocladius subgen. n.
Trissocladius, Zalutschia, Paratrissocladius, and Sublettiella gen. n. (Diptera:
and some related genera (Diptera: Chironomidae). Aquat. Insects 5: 209-226.
Chironomidae). Bull. Fish. Res. Bd Can. Sather, O.A. 1983c. Three new species of
195:287 pp. Lopescladius Oliveira, 1967 (syn. "Cordites"
Sether. O.A. 1977a. Habrobaenus hudsoni n. Brundin, 19-56, n. syn.), with a phylogeny of
gen., n. sp. and the immatures of Baeoctenus the Parakiefferiella grotp. Mem. Am. ent.
bicolor Saether (Diptera: Chironomidae). J. Soc. 34: 279-298.
Fish. Res. Bd Can. 34: 2354-2361. Srther, O.A. 198-5a. Male and female imagines
Saether,O.A. 1977b. Taxonomic studies on of Platysmittia bilyji sp. n. (Diptera:
Chironomidae: Nanocladius, Chironomidae) from Manitoba, Canada.
Pseudochironomus, and the Harnischia Entomologica scand. 15: 527-531.
complex. Bull. Fish. Res. Bd Can. 196:. 143 Saether,O.A. 1985b. Heleniella parva sp. n.
pp. (Diptera: Chironomidae) from South
Sather,O.A. 19'79. New name for Beckiella Carolina and Tennessee, U.S.A.
Saether, 1977 (Diptera: Chironomidae) nec Entomologica scand. 1-5: 532-535.
Beckiella Grandjean, 1964 (Acari: Oribatei). Sether, O.A. 198-5c. Apometriocnemus
Entomologica scand. 10: 31-5. fontinalis n. gen., n. sp. (Diptera:
Srether, O.A. 1980a. New name for Oliveria Chironomidae) from Tennessee, U.S.A.
Saether, 1976 (Diptera: Chironomidae) nec Entomologica scand. 15: -536-539.
Oliveria Sutherland, 196-5 (+Cnidaria: Sather, O.A. 1985d. Redefinition and review of
Anthozoa) with a first record for the Thienemannia Kieffer, 1909 (Diptera:
European continent. Entomologica scand. Chironomidae), with the description of Z.
11: 399-400. pilinucha sp. n. Aquat. Insects 7: III-I3I.
Sether. O.A. 1980b. The females and Srether, O.A. 198-5e. A review of Odontomesa
immatures of Paracricofopus Thienemann Pagast, 1947 (Diptera, Chironomidae,
and Harnisch, 1932, with the description of Prodiamesinae). Spixiana Suppl. (Muench.)
a new species (Diptera: Chironomidae). tt; 15-29.
Aquat. Insects 2: 1,29-145. Srether, O.A. 198-5f. Trichochilus, a new genus
Sather, O.A. 1981a. Doncricotopus bicaudatus of Orthocladiinae (Diptera, Chironomidae).
n. gen., n. sp. (Diptera: Chironomidae, Spixiana Suppl. (Muench.) 11: 31-36.
Orthocladiinae) from the Northwest Sather, O.A. 198-59. The imagines of
Territories, Canada. Entomologica scand. Mesosmittia Brundin, 1956, with description
12: 223-229. of seven new species (Diptera,
Saether, O.A. 1981b. Orthocladiinae Chironomidae). Spixiana Suppl. (Muench.)
(Chironomidae: Diptera) from the British ll:37-54.
West Indies with descriptions of Saether,O.A. 198-5h. Diplosmittia carinata spec.
Antillocladius n. gen., Lipurometriocnemus n. nov. from Michigan (Diptera, Chironomidae).
gen., and Diplosmittia n. gen. Entomologica Spixiana Suppl. (Muench.) 11: -5-5-57.
scand. Suppl. 16: l-46. Srether, O.A. 1985i. A review of the genus
Sether, O.A. 1981c. Compteromesa oconeensis Rheocricotopas Thienemann & Harnisch,
gen. n., sp. n., a new Prodiamesinae 1932, with the description of three new
72 NEARCTIC CHIRONOMIDAE
Soponis, A.R. 1983. Orthocladius and Diamesinae. Proc. U.S. natn. Mus. 115:
(Orthocladius) ferringtoni n. sp., from Kansas 85- 136.
(Diptera: Chironomidae). J. Kans. ent. Soc. Sublette, J.E. L964b- Chironomidae (Diptera)
56:571-571. of Louisiana I. Systematics and immature
Splirck, R. L922-7923. Beitrlige zur Kenntnis stages of some lentic chironomids of
der Chironomidenmetamorphose I-IV. Ent. west-central Louisiana. Tulane Stud. Zool.
Meddr. L4: 32-48 11922); 49-109, 11923). 11:109-150.
Sptirck, R. & A. Thienemann. 1924. Sublette, J.E. 1966. Type specimens of
"Metriocnemus" ampullaceus vat. austriacus. Chironomidae (Diptera) in the U.S. National
Verh. int. Verein. theor. angew. Limnol. 2: Museum. J. Kans. ent. Soc. 39: 580-607.
222-223. Sublette, J.E. 1,967a. Type specimens of
Staeger, C. 1839. Systematisk fortegnelse over Chironomidae (Diptera) in the Canadian
de i Danmark hidtilfundne Diptera. National Collections, Ottawa. J. Kans. ent.
Naturhist. Tidsskr. (1838-1839) 2: 549-600. Soc. 40: 290-331.
Staeger, C. 1845. Gr{nlands antliater. Sublette, J.E. 1,967b. Type specimens of
Naturhist. Tidsskr. (1844-1845) ser. 2,7: 346- Chironomidae (Diptera) in the Cornell
369. University Collection. J. Kans. ent. Soc. 40:
Steffan, A.W. 1965. Plecopteracoluthus downesi 477 -564.
gen. et sp. nov. (Diptera: Chironomidae), a Sublette, J.E. l9'70. Type specimens of
species whose larvae live phoretically on Chironomidae (Dipt.) in the Illinois Natural
larvae of Plecoptera. Can. Ent. 97: History Survey Collection, IJrbana. J. Kans.
t323-t344. ent. Soc. 43:. 44-95.
Steiner, B. t983. Paracricotopus mozleyi n. sp. sublette, J.E. & M.s. Sublette. 1965. Family
from Georgia (Diptera: Chironomidae). Chironomidae. Pp- 142-181.. In; Stone, A.,
Mem. Am. ent. Soc. 34: 329-335. C.W. Sabrosky, W.W. Wirth, R.H. Foote, &
Steiner, J.W. & J.L. Hulbert. 1982. Nimbocera J.R. Coulson (Eds). A catalogue of the
pinderi, a new species (Diptera: Diptera of America North of Mexico. U.S.
Chironomidae) from the Southeastern United Dep. Agric. Handb. No. 276: I-1696.
States. Fla Ent. 65: 228-233. Sublette, J.E. & M.S. Sublette. I971. The
Stone, A. & Wirth, W.W. 1947. On the marine Orthocladiinae (Chironomidae, Dipt.) of
midges of the genus Clunio Haliday California. I. The Cricotopus infuscatus
(Diptera, Tendipedidae). Proc. ent. Soc. group. Ent. News 82:. 85-102.
Wash. 49: 2OL-224. Sublette. J.E. & M.S. Sublette. I974. A review
Stori, R. 1936. FOr faunan nya nematocerer. of the genus Chironomus (Diptera,
Notul. ent. 16: 124-125. Chironomidae) V. The maturus complex. -
Stor6, R. 1939. Mitteilungen riber die Stud. nat. Sci. (Portales, NM) 1, 8: 41 pp.
Nematoceren Finnlands II. Notul. ent. 19: Sublette, J.E. & M.S. Sublette. L979. Part II.
16-30. A synopsis of the Chironomidae of New
Strenzke, K. 194I- Terrestrische Chironomiden Mexico. Pp. 54-128. In: Sublette, J.E. (Ed.).
X. Georthocladius luteicornls Goetgh. (mit Evaulation of long term effects of thermal
einem Beitrag von Dr. M. Goetghebuer). effluents on stream biota. Final Rep. N.
Zool. Anz. 132: 177 -185. Mex. Energy Inst. 32: 1-156.
Strenzke, K. 1950. Systematik, Morphologie Sublette, J.E. & M.S. Sublette. 1983. Asheum,
und Okologie der terrestrischen a replacement name in Cbironomidae
Chironomiden. Arch. Hydrobiol. Suppl. 18: (Diptera). Ent. News 94. 34.
207-4r4. Sublette, J.E. & W.W. Wirth- 1972. New
Strenzke, K. & H. Remmert. 1,957. genera and species of West Indian
Terrestrische Chironomiden. XVII. Chironomidae (Diptera). Fla Ent. -55: 1-17.
Thal ass osmittia thalas s ophila (Bequ. u. Srilc, K. & J- ZavreL 7924. O epoikickych a
Goetgh.). Kieler Meeresforsch. I3:, 263-273. parasitickych larvdch Chironomidu. lUber
Sublette, J.E. 1960. Chironomid midges of epoikische und parasitische
California. I. Chironominae, exclusive of Chironomidenlarven). Acta Soc. Sci. nat.
Tanytarsini (=Calopsectrini). Proc. U.S. moravo-siles. 1: 3-53-391.
natn. Mus. 112: L97-226. Thienemann, A. t921. Die Metamorphose der
Sublette, J.E. 1964a. Chironomid midges of Chironomidengattungen Camptocladius,
California. II. Tanypodinae, Podonominae, Dyscamptocladius und Phaenocladias, mit
Bemerkungen tiber die Artdifferenzierung
NEARCTIC CHIRONOMIDAE
bei den Chironomiden riberhaupt. Arch. Tokunaga, M. 7937. Chironomidae from Japan
I{ydrobiol. Suppl. 2: 809-850. (Diptera), IX Tanypodinae and Diamesinae.
Thienemann, A. 1,926. I{ydrobiologische Philipp. J. Sci. 57: 2l-65.
Untersuchungen an den kalten Quellen und Tokunaga, M. 1939. Chironomidae from Japan
Bdcken der Halbinsel Jasmund auf Rtigen. (Diptera). XI. New or little-known midges,
Arch. Hydrobiol. 17: 221-336. with special reference to the metamorphosis
Thienemann, A. 1934. Chironomiden- of torrential species. Philipp. J. Sci. 69:
Metamorphosen. VIII. " Phaenocladius" 297 -345.
Encycl. ent. Diptera 7: 29-46. Tottenham, C.E. 1949. New generic names in
Thienemann,A. 1935. Chironomiden- Staphylinidae (Coeloptera). Proc. R. ent.
Metamorphosen. X. "Orthocladius Soc. Lond. 18: 78.
-Dactylocladizs" (Dipt.). Stettin. ent. Ztg Townes, H.K. 1945. The Nearctic species of
96: 2OI-224. Tendipedini (Diptera, Tendipedidae
Tbienemann, A. 1936a. Alpine Chironomiden. (=Chironomidae)). Am. Midl. Nat. 34:
Ergebnisse von lJntersuchungen in der t-206.
Gegend von Garmisch-Partenkirchen, Verrall, G.H. 79t2. Another hundred new
Oberbayern. Arch. Hydrobiol. 30: L67-252. British species of Diptera. Entomologist's
Thienemann, A. 1936b. Chironomiden- mon. Mag. 48: 20-27, 56-59, 1.44-147;
Metamorphosen. XI. Gattung Eukiefferiella. 190-197.
Stettin. ent. Ztg 9'7: 43-65. Verrill, A.E. 1873. Report upon the
Thienemann, A. 7939. Dritter Beitrag zur invertebrate animals of Vineyard Sound and
Kenntnis der Podonominae (Dipt. the adjacent waters, with an account of the
Chironomidae). (Chironomiden aus physical characters of the region. Pp.295-
Lappland VI). Zool. Anz. 728: 761-176. 178. In: Baird, S.F., Report on the
Thienemann, A. 7947. Lappliindische condition of the sea fisheries of the south
Chironomiden und ihre WohngewAsser. coast of New England in 1871 and 7872.
(Ergebnisse von lJntersuchungen im U.S. Comn. Fish Fisheries. 1: 1-852.
Abiskogebiet in Schwedisch-Lappland). Walker, F. 1848. List of the specimens of
Arch. Hydrobiol. Suppl. 77:- 7-253. dipterous insects in the collection of the
Thienemann, A. 7942. Trichocladius-Arten aus British Museum. Part 1: l-229. London.
dem Lunzer Seen. (Chironomiden aus dem Walker, F. 1856a. Insecta Britannica: Diptera.
Lunzer Seengebiet. V.). Arch. Hydrobiol. 3: l-352. London.
39:294-315. Walker, F. 1856b. Diptera. 1. Pp. 415-474.
Thienemann, A. & O. I{arnisch. 1932. In. Saunders, W.W. (Ed.). 1850-1869.
Chironomiden-Metamorphosen. IV. Die Insecta Saundersiana: Or characters of
Gattung Cricotopus v.d.W. Zool. Anz. 99: undescribed insects in the collection of
13-5- 143. William Wilson Saunders, Esq., F.R.S.,
Thienemann, A. & J.J. Kieffer. 1916. F.L.S., &c. 4 vols. London.
Schwedische Chironomiden. Arch. Walley, G.S. 1925. New Canadian
llydrobiol. Planktonk. Suppl. 2: 483-554. Chironomidae of the genus Tanypus (Dipt.).
Thienemann, A. & F. Kriiger. 1,937. Can. Ent. 57: 271-278.
"Orthocladius" abiskoensis Edwards und Walley, G.S. 1926a. New Canadian
rubicundus Mg., zwei "Puppen-Species" der Chironomidae. Can. Ent. 58: 64-65.
Chironomiden. (Chironomiden aus Lappland Walley, G.S. 1926b. Four new Canadian
II.) Zool. Anz. 1I7: 257-267. Chironomidae. Can. Ent. 58: 205-207.
Thienemann, A. & F. Krriger. 1939- Walley, G.S. 7928a. A new species of
Terrestrische Chironomiden II. Zool. Anz. Cricotopus with a key to the genus (Dipt.,
127: 246-258. Chironomidae). Can. Ent. 6O: 2l-22.
Tilbury,M.R. 1913. Notes on the feeding and Walley, G.S. 1928b. The genus Tanypus in
rearing of the midge, Chironomus cayugae Canada, with a key to the North American
Johannsen. Jl N.Y. ent. Soc. 21: 305-308. species (Diptera, Chironomidae). Ann. ent.
Tokunaga, M. 1933. Chironomidae from Japan Soc. Am. 2L: 581-593.
(Diptera). I. Clunioninae. Philipp. J. Sci. Webb, D.W. 1969. New species of chironomids
5l: 87 -99. from Costello Lake, Ontario (Diptera:
Tokunaga, M. 1936. Chironomidae from Japan Chironomidae). J. Kans. ent. Soc. 42:
(Diptera), VI Diamesinae. Philipp. J. Sci. 91-108.
59: 526-552.
CHIRONOMIDAE DE LA REGION NEARCNQUE
Wiedemann, C.R.W. 1817. Neue Zweiflrigler und Verbreitung. Arch. Hydrobiol. Suppl.
(Diptera Linn.) aus der Gegend um Kiel. 24: l-66.
Wiedemanns zool. Mag. 1: 61-86. WLilker, W. 1959. Drei neue
Wiedemann, C.R.W. 1828. Aussereuropiiische Chironomiden-Arten (Dipt.) und ihre
zweifltigelige Insekten. 1 lIamm, 32 + 608. Bedeutung fr-ir das Konvergenzproblem bei
Wiederholm, T. I975. Description of Imagines und Puppen. Arch. Hydrobiol.
Protanypus saetheri n. sp. from Alaska Suppl.25: 44-64.
(Diptera: Chironomidae). Entomologica Wtilker W.F. & M.G. Butler. 1983.
scand. 6: 224-228. Karyosystematics and morphology of
Wiederholm, T.(Ed.) 1983. Chironomidae of Northern Chironomus (Diptera:
the Holarctic Region. Keys and diagnoses. Chironomidae): Freshwater species with
Part 1 Larvae. Entomologica scand. larvae of tbe salinarius-type. Entomologica
Suppl. -19:- l-437. scand. 14: 127-1.36.
Wiederholm, T.(Ed.) 1986. Chironomidae of Wulp, F.M. van der. 1858. Beschrijving van
the Holarctic Region. Keys and diagnoses. eenige nieuwe of twijfelachtige soorten van
Part 2 Pupae. E,ntomologica scand. Diptera uit de familie der Nemocera.
Suppl. -28: l-482. Tijdschr. Ent. 2: 159-185.
Wiederholm, T.(Ed.) 1989. Chironomidae of Wulp, F.M. van der. 1861. Over Nieuwe
the Holarctic Region. Keys and diagnoses. soorten van Diptera. Tijdschr. Ent. 4: 1-16.
Part 3 Adult males. Entomologica scand. Wutp, F.M. van der. 1867. Eenige Noord-
Suppl. -34: l-532. Americaansche Diptera. Tijdschr. Ent. 12;
Willassen, E. 1985. A review of Diamesa davisi t25-764.
Edwards and the davisi group (Diptera, Wulp, F.M. van der. 1874. Dipterologische
Chironomidae). Spixiana Suppl. (Muench.) aanteekneningen. Tijdschr. Ent. I7: 1.04-
17:109-137. 148.
Williston, S.W. 1896. On the Diptera of St. Zavrel, I. L937. Eine neue Trissocladius art.
Vincent (West Indies). Trans. Roy. Spisy vyddv. prir. Fak. Masaryk. Univ. 239:.
Entomol. Soc. London. III: 253-446. 1.-1.2.
Winnertz, J. 1846. Beschreibung einiger neuer Zavrel, J.1941. Chironomidarum larvae et
Gattungen aus der Ordnung der Zweiflrigler. nymphae llI. ("Pseudokiefferiella"). Ent.
Stettin. ent. Ztg 7: ll-20. Listy 4: 9-14.
Winnertz, J. 1,852. Dipterologisches. Zavrel, J. & F. Pagast. 193-5. Zwei neue
Stettin. ent. ZtE i3: 49-58. Orthocladiinen-Arten aus Hoher Tatra. Cas.
Wirth, W.W. 1949. A revision of the clunionine csl. Spol. ent.32: 156-160.
midges with descriptions of a new genus and Zetterstedt, J.W. 1838. Diptera. Insecta
four new species (Diptera: Tendipedidae). Lapponica 3: 477-868. Leipzig.
Univ. Calif. Publs Ent. 8: 151-182. Zetterstedt, J.W. 18-50. Diptera Scandinaviae.
Wirth, W.W. t957. The species of Cricotopus Diposita et descripta. 9: 3367-3710.
midges living in the blue-green alga Nostoc Zetterstedt, J.W. 1860. Diptera Scandinaviae.
in California (Diptera: Tendipedidae). Diposita et descripta. 14: 6191-6609.
Pan-Pacif. Ent. 33: 12I-126.
Wirth, W.W. & J.E. Sublette. t970. A review
of the Podominae of North America with
descriptions of three new species of
Trichotanypus (Diptera: Chironomidae). J.
Kans. ent. Soc. 43: 33-5-354.
Wrubleski, D.A. & S.S. Roback. 1987. Two
species of Procladius (Diptera:
Chironomidae) from a northern prairie
marsh: descriptions, phenologies and mating
behaviour. J. N. Am. Benthol. Soc. 6: 198-
2r2.
Wrilker, W. 1956. Zur Kenntnis der Gattung
Psectrocladias Kieff. (Dipt., Chironom.).
Individuelle Variabilitdt. Grenzen und
Mdglichkeiten der Artentrennung, Okologie
76 NEARCTIC CHIRONOMIDAE
N o
naelis Roback, Coelotanypus 8 obediens (Johannsen), Phaenopsectra 50
naevus (Mitchell), Stictochironomus 54 oblidens (Walker), Orthocladius 3L
nais (Townes), Paracladopelma 50 obreptus (Townes), Demeijerea 45
Nanocladius Kieffer 30 obriaini Hayes & Murray, Tokunagaia 39
nansenl Kieffer, Orthocladius 31 obscurifuscus Sublette & Sublette,
nanseni (Kieffer), Pseudosmittia a! Cricotopus 25
nanseni (Kieffer), Smittia J6 obscuripes (Holmgren), Metriocnemus 30
nanus (Llndbeck), Pa rametriocnemus J4 obscurus (Johannsen), Cardiocladius 2l
narda Roback, Krenopelopia 1,2 obsepta (Webb), Zalutschia 40
nasthecus Sather, Paraphaenocladius 34 obtusilobus (Malloch), Cryptochironomus 44
Natarsia Fittkau 10 obtusum Townes, Polypedilum )l
NATARSIINI 10 obumbratus Johannsen, Orthocladius 3L
nalchitocheae (Sublette), Einfeldia 46 obvius (Walker), Psectrocladius 35
natvigi (Goetghebuer), Hydrobaenus 27 occidentalis (Coquillett), Prodiamesa 20
nalvigi (Goetghebuer), Paratanytarsus 57 oceanicus (Packard), Cricotopus 25
nearctic a Brundin, Lasiodiamesa ,l
ochrealus (Townes), Chironomus
nearctica Sether, Odontomesa 19 oconeensis Sether, Compteromesa . t9
needhamii (Johannsen), Polypedilum 52 octomerus Srether, Hydrobaenus 28
nemorosus Edwards, Tanytarsus 59 octopunctatus (Loew), Polypedilum 52
neofl avellus Malloch, Tanytarsus 59 Odontomesa Pagast 19
neomodestus (Malloch), Dicrotendipes 46 Oeklandia Kieffer 56
neopilosella Beck & Beck, Labrundinia . . . . L2 ogemawi Roback, Trissopelopia 74
neopunctipennis Sublette, Tanypus I6 okoboji (Walley), Telopelopia I4
nephopterus (Mitchell), Srenochironomus 53 oldenbergi Goetghebuer, Endochironomus 46
nereis (Townes), Paracladopelma 50 olivacea (Meigen), Prodiamesa 20
nerius (Curran), Compterosmittia zz oliveri Se,ther, Chaetocladius '))
nervosus (Staeger), Dicrotendipes 46 oliveri Srether, Heterotrissocladius 27
netta Townes, Pseudochironomus 55 oliverl Soponis, Orthocladius
nietus Roback, Procladius 15 Oliveria Sather JI
niger Andersen, Tanytarsus 59 Oliveridia Sather JI
niger Borkent, Stenochironomus )5 olivetus Boesel, Cricotopus 'r<
nigra Brundin, Parakiefferiella . . J5 Omisus Townes 49
nigra Roback, Ablabesmyia 10 ontario (Walley), Polypedilum 51
nigricans (Johannsen), Endochironomus 46 onteona Roback, Diamesa I7
nigripila (Johannsen), Micropsectra 56 ophioides Townes, Polypedilum 52
nigrilibla Walker, Chironomus 43 orbicus (Townes), Chernovskiia 4l
nigriturn Townes, Polypedilum fl Oreadomyia Kevan & Cutten-Ali-Khan 3l
nigri(us Malloch, Orthocladius 5Z ornata Beck & Beck, Ablabesmyia 10
nigrohalterale (Malloch), ornatipes (Kieffer), Stictochironomus <A
vockerothlHansen,Diamesa ..... 18
vockerothl Rasmussen, Chironomus . . . . . . 42
vockerothiSrether,Zalutschia .... 40
vulneratus (Zetterstedt),
Demicryptochironomus ...... 45
w
walleyiTownes,Polypedilum ..... 51
waltliiMeigen, (Diamesa) ....... 19
wartinbei(Roback),Reomyia .... 13
whltseli Sublette & Sublette, Chironomus . . 43
wienslSrether, Orthocladius .....32
wilhmiRoback,Procladius ...... 15
wingoi Sather & Sublette, Pseudorthocladius 36
winnelll Jackson, Paracladopelma . . . . 50
wirlhi Roback, Clinotanypus 8
wirlhl Sather & Sublette, Georthocladius . . 26
WlrthiellaSublette .....48
woodi Borkent, Stenochironomus . . . . . . . . 53
x
xantha(Roback),Micropsectra ...57
xanthusSublette,Tanytarsus ..,.. 59
xena (Roback), Thienemanniella . . . . . . . . 39
XenochironomusKieffer ........ 54
xenolabis (Kieffer), Xenochironomus . . . . . 54
XenopelopiaFittkau ....14
Xestochironomus Sublette & Wirth . . . . . . 54
xcthis (Roback), Zalutschia . . . . . . . . . 40
XylolopusOliver ......39
Y
yaquina Cranston & Judd, Metriocnemus 30
z
ZalutschiaLipina ......40
zalulschicolaLipina,Zalutschia ... 40
zavrellPagast,Sympotthastia ..... 19
Zavrelia Kieffer .......59
ZavrellellaKieffer ......55
ZavrelimyiaFittkau .....14
zetterstedtlBrundin, Psectrocladius ... 35
zonopterus (Mitchell), Stenochironomus 53